Was the purpose of these holes ever explained? I looked through the whole thread again and didn't see an explanation of these holes but there is a strong possibility that I missed the explanation.
primarily for the reason snook mentions but also to break the water surface tension in the res below in order to minimize surface precipitates which tend to form on top of the protein layer.
in any closed water system containing life forms there will form a protein layer formed by animal wastes or plant exudate.
in the ppk the tailpiece by itself does not break the tension as it ends below the surface.
another way to do it would be to use a small air stone or diffuser in each plant res.