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So we could be looking at something that acts dominate in one situation then can act recessive? So we know from Epistasis that it might act antagonist then be complimentary like it can go either or is just in balance. We know that the male can be selfed and then females grown out and each on can be observed with others and frequencies can be tabulated. Some form of marker can be found to measure gene distance for traits that are associated with hermi or non hermi characteristics. Just spitballing some of the things that are being thrown around. Maybe we take into account that the frequency of traits can be correlated with cross over with chromosomes and the fact that some traits are expressed randomly but still have to interact with non linked chromosomes also experiencing cross over making mapping of two chromosomes at same time or maybe three and acting complimentary in on situations and antagonist in others. Not sure how you could map the phenotypes for multiple chromosomes experiencing cross over at same time and trying to figure non chromosome linked Epistasis. I know it is mixed up but like I said spitballing stuff maybe we all figure it out tomorrow maybe not